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Aria, Emily, Spencer, Hanna and Alison have been best friends since the third grade. Each goes everywhere together, considering no-one will come between them. If anyone is the ringleader of the group it is Alison, and the other young ladies cannot help but confide all their secrets to her.One night time, during a sleepover, Alison runs missing. Her person is never found. Girls mourn her fatality but move aside after time, assuming their secrets have vanished with Alison too.3 years later and Aria is having an affair with her educator; Emily is questioning her sexuality; Hanna is a thief; and Spencer is flirting with her sister's fiance. They all think their secrets are safe, until they starting acquiring messages from the strange A - who has learned exactly what they are all up to, and is also threatening to spill the beans . . .